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cellule procaryote autre que les bactéries, pourvue d'une membrane sans acide gras et d'une paroi de pseudopeptidoglycane. Son chromosome est circulaire mais associé à des histones et lu par des mécanismes de transcription et de traduction similaires aux Eucaryotes. Les métabolismes des Archae sont très variés comme les bactéries, avec en plus la méthanogénèse. On les a d'abord découverts dans les milieux extrêmes mais on les trouve depuis dans tous les biotopes. On ne connait pas d'Archae pathogènes.Les Archae forment un 3eme règne avec les bactéries et les eucaryotes et sont plus proches en termes d'évolution des eucaryotes que des bactéries.

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